Primary and secondary production

Ecosystems (Interaction and interdependence) · Transfers of energy and matter · note 6 of 9

Primary and secondary productionSpec C4.2.15, C4.2.16

In short

Primary production is the accumulation of carbon compounds in biomass by autotrophs. Secondary production is the accumulation of carbon compounds in biomass by heterotrophs. Both are measured as mass of carbon per unit area per unit time, usually g m⁻² yr⁻¹. Secondary production is lower than primary production because carbon compounds are lost as carbon dioxide and water in respiration.

Primary production
Accumulation of carbon compounds in biomass by autotrophs.
Secondary production
Accumulation of carbon compounds in biomass by heterotrophs.

Biomass accumulates when organisms grow or reproduce. Production is a rate, so its units are mass of carbon per unit area per unit time, usually g m⁻² yr⁻¹.

Biomes vary in their capacity to accumulate biomass. Production is high in tropical rainforests, where it is warm, wet and light all year, and low in deserts and tundra, where water or temperature limits photosynthesis and the growing season is short.

Secondary production is lower than primary production in an ecosystem. When carbon compounds are oxidised in cell respiration, they are converted to carbon dioxide and water and leave the organism, so that biomass is lost. Only the part of the food that is not respired, egested or excreted is built into new heterotroph biomass.

Common mistake:

Production is not the same as biomass. Biomass is the mass present at one moment (g m⁻²); production is how fast it accumulates (g m⁻² yr⁻¹).

Written and checked against the IB Biology SL specification · Updated October 2026

Frequently asked questions

Why is energy lost between trophic levels?

Energy is lost between trophic levels because not all of an organism is eaten, not all food eaten is digested, and much of the absorbed energy is converted to heat in cell respiration. Heat is lost to the environment. Typically only about 10% of the energy passes to the next level, which limits food chain length.

What is the difference between an autotroph and a heterotroph?

An autotroph uses an external energy source, light or oxidation reactions, to synthesise carbon compounds from simple inorganic substances such as carbon dioxide. A heterotroph uses carbon compounds obtained from other organisms, digesting and assimilating them to build the carbon compounds it needs. Both release energy by cell respiration.

Why can matter be recycled in ecosystems but energy cannot?

Matter is recycled because atoms are conserved: decomposers break down dead organic matter and release elements in inorganic forms that producers reuse. Energy cannot be recycled because it is eventually converted to heat by cell respiration, and organisms cannot convert heat back into chemical energy. New energy must keep arriving as sunlight.

All 5 questions on Transfers of energy and matter